Looks like very subtle retouching done. Probably some curves, hue/sat etc.
But if you notice closely, there is some HDR kinda feel to it. The guy says he never uses HDR. How exactly can this kind of an effect be done? If, let's say, he has used Nik Color Efex or any other plug in, what have they done in them and how?
The sky looks dramatic too, how is that done?
I understand this is very very basic, but I am getting curious.
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